Physaria gordonii |
Physaria grahamii |
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Gordon's bladderpod |
Graham's twinpod |
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Habit | Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | (proximal sometimes petiolate, distal sessile); blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | dense. |
loose, (elongated). |
Flowers | sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | (shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20(–26) per ovary; style (1.5–)2–4(–5) mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | flattened. |
plump, (suborbicular). |
2n | = 12, 32. |
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Physaria gordonii |
Physaria grahamii |
|
Phenology | Flowering Feb–Jul. | Flowering May–Jun. |
Habitat | Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 150-1700 m (500-5600 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
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UT |
Discussion | Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif. Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 640. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |