Physaria gordonii |
Physaria eburniflora |
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Gordon's bladderpod |
Devils Gate twinpod |
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Habit | Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). | Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). |
Stems | several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm. |
simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm. |
Basal leaves | blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire. |
blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed). |
Cauline leaves | (proximal sometimes petiolate, distal sessile); blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate. |
(2–4); blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute). |
Racemes | dense. |
condensed. |
Flowers | sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base). |
sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median); petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm. |
(divaricate-ascending, nearly straight), 6–10 mm. |
Fruits | (shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20(–26) per ovary; style (1.5–)2–4(–5) mm. |
strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery); valves (retaining seeds after dehiscence), pubescent; replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse; ovules 4–8 per ovary; style 4–5 mm, (sparsely pubescent or glabrous). |
Seeds | flattened. |
plump. |
2n | = 12, 32. |
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Physaria gordonii |
Physaria eburniflora |
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Phenology | Flowering Feb–Jul. | Flowering May–Jun. |
Habitat | Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields | Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus |
Elevation | 150-1700 m (500-5600 ft) | 1800-3000 m (5900-9800 ft) |
Distribution |
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
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WY |
Discussion | Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif. Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 640. | FNA vol. 7, p. 635. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia | |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | Rollins: Brittonia 33: 333. (1981) |
Web links |