Physaria gordonii |
Physaria douglasii |
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Gordon's bladderpod |
Columbia bladder-pod, Douglas' bladder-pod |
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Habit | Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | ||||
Stems | several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm. |
simple from base, erect, (usually unbranched), to 4.5 dm. |
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Basal leaves | blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire. |
blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
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Cauline leaves | (proximal sometimes petiolate, distal sessile); blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate. |
similar to basal, blade narrowly linear or, sometimes, orbicular. |
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Racemes | dense. |
loose (lax). |
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Flowers | sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base). |
sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
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Fruiting pedicels | (divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm. |
(recurved, straight, curved, or sigmoid), 6–20 mm. |
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Fruits | (shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20(–26) per ovary; style (1.5–)2–4(–5) mm. |
obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 12, 32. |
= 10, 30. |
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Physaria gordonii |
Physaria douglasii |
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Phenology | Flowering Feb–Jul. | |||||
Habitat | Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields | |||||
Elevation | 150-1700 m (500-5600 ft) | |||||
Distribution |
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
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ID; MT; OR; WA; BC
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Discussion | Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif. Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 640. | FNA vol. 7, p. 634. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia | Lesquerella douglasii | ||||
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
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