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Gordon's bladderpod

Dorn's twinpod, tunp range twinpod

Habit Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). Perennials; (compact); caudex simple, (stout); densely (silvery) pubescent throughout (except style), trichomes several-rayed, rays furcate, fused at base, (umbonate, tuberculate throughout).
Stems

several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm.

simple from base, erect, (arising from a condensed rosette), to 1 dm.

Basal leaves

blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire.

(ascending or erect);

blade elliptic to oblanceolate to obovate, (usually curled from middle to apex), (1.5–)5–7 cm (width 12–20 mm), margins entire.

Cauline leaves

(proximal sometimes petiolate, distal sessile);

blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate.

(1–5);

blade oblanceolate, 1–2.5 cm, margins entire.

Racemes

dense.

compact (or elongated in fruit, to 1 dm, barely exceeding leaves).

Flowers

sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base).

sepals (erect), oblong to linear or spatulate, 5.5–7 mm;

petals spatulate, 10–14 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm.

(divaricate-ascending, slightly curved), 7–18 mm.

Fruits

(shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20(–26) per ovary;

style (1.5–)2–4(–5) mm.

didymous, irregular, highly inflated, 8–11(–18) × 10–15 mm, (papery, basal sinus shallower than the deep apical sinus; valves retaining seeds after dehiscence);

replum obovate, not constricted, 1–1.8 mm, apex obtuse, as wide as or wider than fruit;

ovules (4–)8(–12) per ovary;

style 4–6 mm, (glabrous).

Seeds

flattened.

flattened, (oblong to elliptic, thin-margined or not).

2n

= 12, 32.

Physaria gordonii

Physaria dornii

Phenology Flowering Feb–Jul. Flowering May–Jun.
Habitat Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields Calcareous shale, slopes, ridges
Elevation 150-1700 m (500-5600 ft) 1900-2200 m (6200-7200 ft)
Distribution
from FNA
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif.

Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 640. FNA vol. 7, p. 634.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) Lichvar: Brittonia 35: 150, figs. 1–3. (1983)
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