Physaria gordonii |
Physaria engelmannii |
|
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Gordon's bladderpod |
Engelmann's bladderpod |
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Habit | Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). |
Stems | several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm. |
few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
Basal leaves | blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire. |
blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
Cauline leaves | (proximal sometimes petiolate, distal sessile); blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate. |
(proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
Racemes | dense. |
dense, (subumbellate). |
Flowers | sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base). |
sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
Fruiting pedicels | (divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm. |
(ascending), relatively short. |
Fruits | (shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20(–26) per ovary; style (1.5–)2–4(–5) mm. |
(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
Seeds | flattened. |
flattened. |
2n | = 12, 32. |
= 12, 24, 36. |
Physaria gordonii |
Physaria engelmannii |
|
Phenology | Flowering Feb–Jul. | Flowering Apr–May. |
Habitat | Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops |
Elevation | 150-1700 m (500-5600 ft) | 150-400 m (500-1300 ft) |
Distribution |
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
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OK; TX |
Discussion | Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif. Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 640. | FNA vol. 7, p. 635. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |