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Geyer's bladderpod, Geyer's twin-pod

Kodachrome bladderpod

Habit Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). Perennials; caudex (buried), branched, (forming hard mats); densely pubescent, trichomes several-rayed, rays furcate or bifurcate, (tuberculate).
Stems

several from base, decumbent, (arising laterally, unbranched), 1–3 dm.

several from base, erect, (unbranched), 0.2–0.3 dm.

Basal leaves

(numerous); (petiole slender, rarely with a few broad teeth);

blade obovate, 3–7 cm, margins entire.

(few), similar to cauline.

Cauline leaves

blade oblanceolate, 1.5–3 cm, margins entire.

(petiole not differentiated from blade);

blade (somewhat succulent), linear to narrowly oblanceolate, 5–12 mm, margins entire.

Racemes

loose.

dense, (few-flowered).

Flowers

sepals oblong, 5–7 mm;

petals (yellow to purplish), spatulate, 8–12 mm.

sepals (yellowish), elliptic, 3–4.5 mm;

petals (erect or, more commonly, arching), spatulate to oblanceolate, 5.8–7 mm, (claw not or weakly differentiated from blade).

Fruiting pedicels

(ascending or spreading, slightly curved or sigmoid), 1–2 cm.

(ascending to divaricate-ascending, ± straight), 3.5–6 mm.

Fruits

obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading;

replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse;

ovules 4–6 per ovary;

style 5–7 mm.

(coppery or reddish brown in age), broadly ovoid, slightly inflated, 3–4 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

flattened.

Physaria geyeri

Physaria tumulosa

Phenology Flowering May–Jun.
Habitat Barren white knolls surrounded by sagebrush, pinyon pine, and Utah juniper
Elevation 1600-1800 m (5200-5900 ft)
Distribution
from FNA
ID; MT; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria tumulosa is morphologically similar to 55. P. navajoensis of northeastern Arizona and northwestern New Mexico, and differing very subtly. It has been long treated as an infraspecific taxon of P. hitchcockii; unpublished molecular data do not support that disposition. It is found on knolls of the Winsor Member of the Carmel Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Replum apices acute; ovules 4 per ovary; petals yellow, drying yellow.
subsp. geyeri
1. Replum apices obtuse; ovules usually 6 per ovary; petals purplish or light yellow, drying purplish.
subsp. purpurea
Source FNA vol. 7, p. 638. FNA vol. 7, p. 664.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. geyeri subsp. geyeri, P. geyeri subsp. purpurea
Synonyms Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri Lesquerella hitchcockii subsp. tumulosa, Lesquerella tumulosa, P. rubicundula var. tumulosa
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) (Barneby) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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