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Geyer's bladderpod, Geyer's twin-pod

Nelson's bladderpod

Habit Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout).
Stems

several from base, decumbent, (arising laterally, unbranched), 1–3 dm.

few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves).

Basal leaves

(numerous); (petiole slender, rarely with a few broad teeth);

blade obovate, 3–7 cm, margins entire.

blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire.

Cauline leaves

blade oblanceolate, 1.5–3 cm, margins entire.

(absent or few);

similar to basal, blade linear.

Racemes

loose.

dense, (few-flowered).

Flowers

sepals oblong, 5–7 mm;

petals (yellow to purplish), spatulate, 8–12 mm.

sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals (bright yellow), lingulate, 6–8 mm.

Fruiting pedicels

(ascending or spreading, slightly curved or sigmoid), 1–2 cm.

(loosely sigmoid), 3–5 mm.

Fruits

obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading;

replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse;

ovules 4–6 per ovary;

style 5–7 mm.

lanceolate, compressed apically, 2.5–3(–4) mm;

valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside;

ovules 4–8 per ovary;

style 2.5–4 mm.

Seeds

plump, (oblong).

Physaria geyeri

Physaria nelsonii

Phenology Flowering May-mid Jun.
Habitat Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants
Elevation 1600-2300 m (5200-7500 ft)
Distribution
from FNA
ID; MT; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
UT; WY
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Replum apices acute; ovules 4 per ovary; petals yellow, drying yellow.
subsp. geyeri
1. Replum apices obtuse; ovules usually 6 per ovary; petals purplish or light yellow, drying purplish.
subsp. purpurea
Source FNA vol. 7, p. 638. FNA vol. 7, p. 652.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. geyeri subsp. geyeri, P. geyeri subsp. purpurea
Synonyms Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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