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Geyer's bladderpod, Geyer's twin-pod

Navajo bladderpod

Habit Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo).
Stems

several from base, decumbent, (arising laterally, unbranched), 1–3 dm.

several from base (crowded), erect, not exceeding leaves.

Basal leaves

(numerous); (petiole slender, rarely with a few broad teeth);

blade obovate, 3–7 cm, margins entire.

usually absent.

Cauline leaves

blade oblanceolate, 1.5–3 cm, margins entire.

(petiole not differentiated from blade);

blade linear-oblanceolate, 3–8(–13) mm, margins entire.

Racemes

loose.

(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves).

Flowers

sepals oblong, 5–7 mm;

petals (yellow to purplish), spatulate, 8–12 mm.

sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate);

petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle).

Fruiting pedicels

(ascending or spreading, slightly curved or sigmoid), 1–2 cm.

(ascending to divaricate-ascending, straight), 3.5–6 mm.

Fruits

obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading;

replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse;

ovules 4–6 per ovary;

style 5–7 mm.

(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute);

valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not);

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

plump or slightly flattened, (strongly mucilaginous).

Physaria geyeri

Physaria navajoensis

Phenology Flowering May-early Jun.
Habitat Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation 2200-2400 m (7200-7900 ft)
Distribution
from FNA
ID; MT; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Replum apices acute; ovules 4 per ovary; petals yellow, drying yellow.
subsp. geyeri
1. Replum apices obtuse; ovules usually 6 per ovary; petals purplish or light yellow, drying purplish.
subsp. purpurea
Source FNA vol. 7, p. 638. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. geyeri subsp. geyeri, P. geyeri subsp. purpurea
Synonyms Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri Lesquerella navajoensis
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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