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Geyer's bladderpod, Geyer's twin-pod

Fendler bladderpod, Fendler's bladderpod

Habit Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center).
Stems

several from base, decumbent, (arising laterally, unbranched), 1–3 dm.

several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm.

Basal leaves

(numerous); (petiole slender, rarely with a few broad teeth);

blade obovate, 3–7 cm, margins entire.

blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate.

Cauline leaves

blade oblanceolate, 1.5–3 cm, margins entire.

(shortly petiolate);

blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute).

Racemes

loose.

loose to somewhat dense.

Flowers

sepals oblong, 5–7 mm;

petals (yellow to purplish), spatulate, 8–12 mm.

sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate);

petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short).

Fruiting pedicels

(ascending or spreading, slightly curved or sigmoid), 1–2 cm.

(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm.

Fruits

obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading;

replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse;

ovules 4–6 per ovary;

style 5–7 mm.

globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute);

valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout;

replum as wide as or wider than fruit;

ovules (12–)20–32(–40) per ovary;

style (2–)3–6 mm.

Seeds

flattened.

2n

= 12, 14, 24.

Physaria geyeri

Physaria fendleri

Phenology Flowering Mar–May.
Habitat Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas
Elevation 100-2000 m (300-6600 ft)
Distribution
from FNA
ID; MT; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In dry areas, Physaria fendleri may flower following suitable rains at any time of the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Replum apices acute; ovules 4 per ovary; petals yellow, drying yellow.
subsp. geyeri
1. Replum apices obtuse; ovules usually 6 per ovary; petals purplish or light yellow, drying purplish.
subsp. purpurea
Source FNA vol. 7, p. 638. FNA vol. 7, p. 636.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. geyeri subsp. geyeri, P. geyeri subsp. purpurea
Synonyms Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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