Physaria geyeri |
Physaria engelmannii |
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Geyer's bladderpod, Geyer's twin-pod |
Engelmann's bladderpod |
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Habit | Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). | ||||
Stems | several from base, decumbent, (arising laterally, unbranched), 1–3 dm. |
few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
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Basal leaves | (numerous); (petiole slender, rarely with a few broad teeth); blade obovate, 3–7 cm, margins entire. |
blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
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Cauline leaves | blade oblanceolate, 1.5–3 cm, margins entire. |
(proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
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Racemes | loose. |
dense, (subumbellate). |
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Flowers | sepals oblong, 5–7 mm; petals (yellow to purplish), spatulate, 8–12 mm. |
sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
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Fruiting pedicels | (ascending or spreading, slightly curved or sigmoid), 1–2 cm. |
(ascending), relatively short. |
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Fruits | obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading; replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse; ovules 4–6 per ovary; style 5–7 mm. |
(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
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Seeds | flattened. |
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2n | = 12, 24, 36. |
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Physaria geyeri |
Physaria engelmannii |
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Phenology | Flowering Apr–May. | |||||
Habitat | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops | |||||
Elevation | 150-400 m (500-1300 ft) | |||||
Distribution |
ID; MT; OR; WA
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OK; TX |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 638. | FNA vol. 7, p. 635. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | ||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |