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Geyer's bladderpod, Geyer's twin-pod

golden bladderpod

Habit Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). Biennials or perennials; (short-lived); caudex branched; densely pubescent, trichomes (sessile or short-stalked, simple or not), 5–9-rayed, rays furcate, (fine, smooth or finely tuberculate).
Stems

several from base, decumbent, (arising laterally, unbranched), 1–3 dm.

several from base, erect or outer ones decumbent or procumbent, (sometimes much-branched distally), to 6 dm.

Basal leaves

(numerous); (petiole slender, rarely with a few broad teeth);

blade obovate, 3–7 cm, margins entire.

blade obovate or rhombic, to ca. 2.5 cm, margins usually shallowly dentate, sometimes lyrate-pinnatifid.

Cauline leaves

blade oblanceolate, 1.5–3 cm, margins entire.

(proximal shortly petiolate, distal sessile);

blade obovate to rhombic or oblanceolate, 2–4(–6) cm, margins entire or shallowly and remotely dentate.

Racemes

loose.

usually dense, (several-flowered).

Flowers

sepals oblong, 5–7 mm;

petals (yellow to purplish), spatulate, 8–12 mm.

sepals ovate or oblong (tapering at base), 3.6–4.8(–5.3) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals obovate to spatulate, 4.5–7.5 mm, (blade narrowed to broad claw, margins sinuate).

Fruiting pedicels

(ascending or spreading, slightly curved or sigmoid), 1–2 cm.

(strongly recurved), to 20 mm.

Fruits

obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading;

replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse;

ovules 4–6 per ovary;

style 5–7 mm.

(± pendent), ovoid, obcompressed, or globose, compressed, 4–6(–8) mm;

valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules usually 4, rarely 6, per ovary;

style 2.5–3.6 mm.

Seeds

flattened.

2n

= 14.

Physaria geyeri

Physaria aurea

Phenology Flowering Jun–Aug.
Habitat Open sites and bare areas in rocky limestone soil in mountains, roadbanks, open woods
Elevation 2000-2800 m (6600-9200 ft)
Distribution
from FNA
ID; MT; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
NM
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria aurea (known from the Jicarilla and Sacramento mountains) is similar to 35. P. gooddingii, which is found farther west in the mountains of Catron and Sierra counties, New Mexico, and Greenlee County, Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Replum apices acute; ovules 4 per ovary; petals yellow, drying yellow.
subsp. geyeri
1. Replum apices obtuse; ovules usually 6 per ovary; petals purplish or light yellow, drying purplish.
subsp. purpurea
Source FNA vol. 7, p. 638. FNA vol. 7, p. 628.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. geyeri subsp. geyeri, P. geyeri subsp. purpurea
Synonyms Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri Lesquerella aurea
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) (Wooton) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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