Physaria geyeri |
Physaria arizonica |
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Geyer's bladderpod, Geyer's twin-pod |
Arizona bladderpod |
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Habit | Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). | ||||
Stems | several from base, decumbent, (arising laterally, unbranched), 1–3 dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
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Basal leaves | (numerous); (petiole slender, rarely with a few broad teeth); blade obovate, 3–7 cm, margins entire. |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
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Cauline leaves | blade oblanceolate, 1.5–3 cm, margins entire. |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
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Racemes | loose. |
dense, often subcorymbiform. |
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Flowers | sepals oblong, 5–7 mm; petals (yellow to purplish), spatulate, 8–12 mm. |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
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Fruiting pedicels | (ascending or spreading, slightly curved or sigmoid), 1–2 cm. |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
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Fruits | obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading; replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse; ovules 4–6 per ovary; style 5–7 mm. |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
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Seeds | flattened. |
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2n | = 10. |
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Physaria geyeri |
Physaria arizonica |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine | |||||
Elevation | 1000-2200 m (3300-7200 ft) | |||||
Distribution |
ID; MT; OR; WA
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AZ; UT
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 638. | FNA vol. 7, p. 627. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis | ||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |