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point-tip twinpod

Newberry twinpod, Newberry's twinpod

Habit Perennials; caudex branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays mostly furcate, (arms of unequal lengths, finely tuberculate). Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length.
Stems

several from base, erect or lateral decumbent, (unbranched), 1–2 dm.

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

Basal leaves

(petiole usually winged);

blade broadly oblanceolate, 3–8 cm, margins usually dentate or pinnatifid, rarely subentire, (terminal lobe acute or obtuse, not rounded).

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

Cauline leaves

blade spatulate to linear-oblanceolate, 1–3 cm, margins usually entire, rarely toothed, (apex acute).

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

Racemes

loose (and greatly elongated in fruit) to congested.

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

Flowers

sepals linear-oblong, 5–7 mm;

petals spatulate, 9–11 mm.

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

Fruiting pedicels

(recurved), 6–15 mm.

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

Fruits

(usually pendent on arching pedicels, less frequently widely divergent), irregular in shape, (base obtuse or slightly cordate, apex deeply and broadly notched), not strongly inflated, 8–11 × 8–12 mm, (papery);

valves retaining seeds after dehiscence;

replum linear-oblong, constricted, 2.5–4 mm, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–8 mm.

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

Seeds

flattened.

slightly flattened, (ovate).

Physaria floribunda

Physaria newberryi

Distribution
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves inflated, membranous; trichomes appressed.
subsp. floribunda
1. Fruit valves usually not inflated, coriaceous; trichomes slightly spreading.
subsp. osterhoutii
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 637. FNA vol. 7, p. 652.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. floribunda subsp. floribunda, P. floribunda subsp. osterhoutii
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi
Name authority Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861)
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