Physaria fendleri |
Physaria spatulata |
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Fendler bladderpod, Fendler's bladderpod |
alpine bladderpod, spatula-leaf bladderpod |
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Habit | Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). | Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate). |
Stems | several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm. |
simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm. |
Basal leaves | blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate. |
(erect to prostrate, petiole distinct from blade); blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded). |
Cauline leaves | (shortly petiolate); blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute). |
blade spatulate, distinctly different from basal. |
Racemes | loose to somewhat dense. |
moderately dense, (6–20-flowered). |
Flowers | sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate); petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short). |
sepals (pale yellow), elliptic, 3.5–5 mm; petals lingulate, 6–9 mm. |
Fruiting pedicels | (divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm. |
(strongly sigmoid), 10–20 mm (2 times longer than fruits). |
Fruits | globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute); valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout; replum as wide as or wider than fruit; ovules (12–)20–32(–40) per ovary; style (2–)3–6 mm. |
lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed); valves pubescent, trichomes sparse and closely appressed to surface; ovules 4–8 per ovary; style 2.5–6 mm (usually ± equal in length to mature fruit). |
Seeds | flattened. |
plump. |
2n | = 12, 14, 24. |
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Physaria fendleri |
Physaria spatulata |
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Phenology | Flowering Mar–May. | Flowering May-early Jul. |
Habitat | Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas | Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates |
Elevation | 100-2000 m (300-6600 ft) | 900-2900 m (3000-9500 ft) |
Distribution |
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
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MT; ND; NE; SD; WY; AB; SK
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Discussion | In dry areas, Physaria fendleri may flower following suitable rains at any time of the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 636. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera | Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (Rydberg) Grady & O’Kane: Novon 17: 190. (2007) |
Web links |