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Fendler bladderpod, Fendler's bladderpod

Hitchcock's bladderpod

Habit Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate).
Stems

several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm.

few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm.

Basal leaves

blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate.

(petiole and blade differentiated or not);

blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire.

Cauline leaves

(shortly petiolate);

blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute).

similar to basal, smaller.

Racemes

loose to somewhat dense.

dense.

Flowers

sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate);

petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short).

sepals narrowly lanceolate to lanceolate, 2.8–6 mm;

petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm.

(ascending, straight or slightly curved), 2–6 mm.

Fruits

globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute);

valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout;

replum as wide as or wider than fruit;

ovules (12–)20–32(–40) per ovary;

style (2–)3–6 mm.

(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute);

valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.7–6 mm.

Seeds

flattened.

flattened.

2n

= 12, 14, 24.

Physaria fendleri

Physaria hitchcockii

Phenology Flowering Mar–May.
Habitat Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas
Elevation 100-2000 m (300-6600 ft)
Distribution
from FNA
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

In dry areas, Physaria fendleri may flower following suitable rains at any time of the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation).
subsp. rubicundula
1. Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah
→ 2
2. Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada.
subsp. hitchcockii
2. Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada.
subsp. confluens
Source FNA vol. 7, p. 636. FNA vol. 7, p. 643.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hitchcockii subsp. confluens, P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
Synonyms Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera Lesquerella hitchcockii
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002)
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