Physaria fendleri |
Physaria hitchcockii |
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Fendler bladderpod, Fendler's bladderpod |
Hitchcock's bladderpod |
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Habit | Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
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Basal leaves | blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate. |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
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Cauline leaves | (shortly petiolate); blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute). |
similar to basal, smaller. |
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Racemes | loose to somewhat dense. |
dense. |
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Flowers | sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate); petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short). |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm. |
(ascending, straight or slightly curved), 2–6 mm. |
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Fruits | globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute); valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout; replum as wide as or wider than fruit; ovules (12–)20–32(–40) per ovary; style (2–)3–6 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 12, 14, 24. |
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Physaria fendleri |
Physaria hitchcockii |
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Phenology | Flowering Mar–May. | |||||||||
Habitat | Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas | |||||||||
Elevation | 100-2000 m (300-6600 ft) | |||||||||
Distribution |
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
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NV; UT |
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Discussion | In dry areas, Physaria fendleri may flower following suitable rains at any time of the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 636. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera | Lesquerella hitchcockii | ||||||||
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |