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Engelmann's bladderpod

strong bladderpod

Habit Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). Perennials; caudex branched, (thickened); densely pubescent, trichomes (sessile), several-rayed, rays furcate or bifurcate, usually fused toward base, (strongly tuberculate).
Stems

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

several from base, erect or outer decumbent, (unbranched), to 2 dm.

Basal leaves

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

blade elliptic to lanceolate or obovate, 3–8 cm, margins entire.

Cauline leaves

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

(proximal shortly petiolate or sessile, distal sessile);

blade elliptic or obovate, to 2 cm, margins entire.

Racemes

dense, (subumbellate).

dense.

Flowers

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

sepals narrowly elliptic or oblong, 4.5–5.3 mm, (tapering to the somewhat thickened, cucullate apex, lateral pair subsaccate);

petals (bright yellow), lingulate or broadly obovate, 7.5–8.5 mm, (narrowing to broad claw, joining in an arch, margins lacerate).

Fruiting pedicels

(ascending), relatively short.

(divaricate-ascending to horizontal, straight to loosely curved), to 15 mm.

Fruits

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

(sessile or substipitate), suborbicular to broadly ovate or ellipsoid, slightly compressed, 6–8 mm;

valves pubescent;

ovules 12–22 per ovary;

style 2–3 mm.

Seeds

flattened.

flattened.

2n

= 12, 24, 36.

= 10.

Physaria engelmannii

Physaria valida

Phenology Flowering Apr–May. Flowering Apr–May.
Habitat Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops Limestone soils, steep slopes, roadcuts, open woods
Elevation 150-400 m (500-1300 ft) 1900-2200 m (6200-7200 ft)
Distribution
from FNA
OK; TX
[BONAP county map]
from FNA
NM; TX
[BONAP county map]
Discussion

Physaria valida is known from the Sacramento and White mountains of south central New Mexico, and southward through the Guadalupe Mountains to Hudspeth County, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 635. FNA vol. 7, p. 664.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. vicina, P. vitulifera
Synonyms Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella Lesquerella valida, Lesquerella lepidota
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Greene) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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