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Engelmann's bladderpod

mountain-view bladderpod

Habit Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). Perennials; caudex branched, (densely cespitose and forming hemispheric mounds); densely pubescent, trichomes 5-rayed, rays bifurcate near base, fused at base, (strongly tuberculate throughout).
Stems

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

few to several from base, erect, (usually exceeding basal leaves), 0.3–0.7 dm.

Basal leaves

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

blade linear-spatulate, 1.5–4 cm, (base narrowed gradually to petiole), margins entire.

Cauline leaves

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

blade spatulate, similar to basal.

Racemes

dense, (subumbellate).

crowded in distal 1/3, (4–10-flowered).

Flowers

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

sepals (pale yellow), oblong to elliptic, 3–4 mm, (median pair usually thickened apically, cucullate);

petals (sometimes with slight tinge of orange basally), lingulate, 4–6 mm.

Fruiting pedicels

(ascending), relatively short.

(loosely to strongly sigmoid), 6–10 mm.

Fruits

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

ellipsoid, slightly inflated (somewhat latiseptate), 4–5 mm, (apex acute);

valves pubescent, trichomes erect, appearing slightly shaggy;

ovules 4–8 per ovary;

styles 2.5–3 mm, (shorter than mature fruits).

Seeds

flattened.

± flattened, convex on outer side.

2n

= 12, 24, 36.

Physaria engelmannii

Physaria pycnantha

Phenology Flowering Apr–May. Flowering late May–Jun(-Jul).
Habitat Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops Dry, windswept knolls of limestone gravel, with other cushion-forming plants
Elevation 150-400 m (500-1300 ft) 1600-2300 m (5200-7500 ft)
Distribution
from FNA
OK; TX
[BONAP county map]
from FNA
ID; MT
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria pycnantha is morphologically similar to 56. P. nelsonii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 635. FNA vol. 7, p. 659.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Grady & O’Kane: Novon 17: 188, fig. 5. (2007)
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