Physaria engelmannii |
Physaria occidentalis |
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Engelmann's bladderpod |
western bladder-pod |
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Habit | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). | Perennials; caudex simple or branched, (thickened, sometimes subterranean); densely pubescent, trichomes (sessile or short-stalked), 5–7-rayed, raysdistinct, bifurcate, (moderately to prominently tuberculate). | ||||
Stems | few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
few to several from base, prostrate to decumbent or erect, (usually unbranched), 0.3–1.5(–3) dm. |
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Basal leaves | blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
(petiole slender); blade suborbicular to obovate or elliptic, 1–8 cm, (base narrowing gradually or abruptly to petiole), margins sinuate-dentate or entire. |
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Cauline leaves | (proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
(proximal shortly petiolate, distal sessile); blade oblanceolate, 0.5–1.5(–2.5) cm, margins entire or remotely dentate. |
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Racemes | dense, (subumbellate). |
dense or loose. |
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Flowers | sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
sepals elliptic or oblong-elliptic, 4.5–7 mm, (median pair cucullate); petals spatulate, 7–9(–14) mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (ascending), relatively short. |
(sigmoid or curved), 5–10(–15) mm. |
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Fruits | (shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
(erect), ellipsoid to obovoid, compressed at apex and sometimes margins (strongly latiseptate), (5–)6–9 mm, (apex acute, often beaked); valves densely pubescent, trichomes appressed or spreading, sometimes sparsely pubescent inside, trichomes 4- or 5-rayed; ovules 4–12 per ovary; style (2–)3–6.5 mm, (often sparsely pubescent). |
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Seeds | flattened. |
inner surface flattened, outer convex. |
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2n | = 12, 24, 36. |
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Physaria engelmannii |
Physaria occidentalis |
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Phenology | Flowering Apr–May. | |||||
Habitat | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops | |||||
Elevation | 150-400 m (500-1300 ft) | |||||
Distribution |
OK; TX |
CA; ID; NV; OR; UT; WA
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 635. | FNA vol. 7, p. 654. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | Vesicaria occidentalis, Lesquerella occidentalis | ||||
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |