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Engelmann's bladderpod

western bladder-pod

Habit Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). Perennials; caudex simple or branched, (thickened, sometimes subterranean); densely pubescent, trichomes (sessile or short-stalked), 5–7-rayed, raysdistinct, bifurcate, (moderately to prominently tuberculate).
Stems

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

few to several from base, prostrate to decumbent or erect, (usually unbranched), 0.3–1.5(–3) dm.

Basal leaves

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

(petiole slender);

blade suborbicular to obovate or elliptic, 1–8 cm, (base narrowing gradually or abruptly to petiole), margins sinuate-dentate or entire.

Cauline leaves

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

(proximal shortly petiolate, distal sessile);

blade oblanceolate, 0.5–1.5(–2.5) cm, margins entire or remotely dentate.

Racemes

dense, (subumbellate).

dense or loose.

Flowers

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

sepals elliptic or oblong-elliptic, 4.5–7 mm, (median pair cucullate);

petals spatulate, 7–9(–14) mm, (claw undifferentiated from blade).

Fruiting pedicels

(ascending), relatively short.

(sigmoid or curved), 5–10(–15) mm.

Fruits

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

(erect), ellipsoid to obovoid, compressed at apex and sometimes margins (strongly latiseptate), (5–)6–9 mm, (apex acute, often beaked);

valves densely pubescent, trichomes appressed or spreading, sometimes sparsely pubescent inside, trichomes 4- or 5-rayed;

ovules 4–12 per ovary;

style (2–)3–6.5 mm, (often sparsely pubescent).

Seeds

flattened.

inner surface flattened, outer convex.

2n

= 12, 24, 36.

Physaria engelmannii

Physaria occidentalis

Phenology Flowering Apr–May.
Habitat Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops
Elevation 150-400 m (500-1300 ft)
Distribution
from FNA
OK; TX
[BONAP county map]
from FNA
CA; ID; NV; OR; UT; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect or decumbent; fruit margins and apices compressed, apices often beaked; valves pubescent inside; ovules 4-8 per ovary; styles frequently curved; n California, c Idaho, nw Nevada, Oregon, Washington.
subsp. occidentalis
1. Stems prostrate to decumbent; fruit, margins not conspicuously compressed, apices somewhat compressed, usually not conspicuously beaked; valves usually glabrous, sometimes sparsely pubescent inside; ovules 8-12 per ovary; styles usually straight; Nevada, w Utah.
subsp. cinerascens
Source FNA vol. 7, p. 635. FNA vol. 7, p. 654.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. occidentalis subsp. cinerascens, P. occidentalis subsp. occidentalis
Synonyms Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella Vesicaria occidentalis, Lesquerella occidentalis
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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