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Engelmann's bladderpod

Middle Butte bladderpod

Habit Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). Perennials; caudex simple; densely pubescent, trichomes (appressed in layers), 5–7-rayed, rays usually bifurcate, sometimes furcate, (thickened toward center).
Stems

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

simple from base, prostrate to decumbent, (unbranched, from within and below leaf clusters, slender), 0.2–0.8 dm.

Basal leaves

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

(tufted, erect or ascending, silvery);

blade linear to oblanceolate or (outer) oblanceolate to obovate or rhombic, 1.5–3.3 cm, (base sometimes subhastate), margins entire or dentate, (often involute).

Cauline leaves

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

blade nearly linear, to 1.5 cm, margins entire.

Racemes

dense, (subumbellate).

dense, (subcorymbose).

Flowers

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

sepals (yellow-green), lanceolate, (2.5–)3.5–4.5 mm;

petals spatulate to oblanceolate, 4–6.5 mm.

Fruiting pedicels

(ascending), relatively short.

(recurved), 5–8 mm.

Fruits

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

(usually pendent), obdeltate, compressed (angustiseptate), 2–4 mm, (wider than long, base tapered to acute angle, apex truncate with inflated shoulders);

valves densely pubescent; (septum obsolete or with a narrow fringe inside of replum, funicles attached close to replum apex);

ovules 4 per ovary;

style 2–4 mm, (slender).

Seeds

flattened.

plump, (mucilaginous when wetted).

2n

= 12, 24, 36.

Physaria engelmannii

Physaria obdeltata

Phenology Flowering Apr–May. Flowering Jun(-Jul).
Habitat Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops Clayey, silty, or gravelly soils, overlaying basalt lava flows, silty playas, sagebrush, barren areas
Elevation 150-400 m (500-1300 ft) 1300-1700 m (4300-5600 ft)
Distribution
from FNA
OK; TX
[BONAP county map]
from FNA
ID
[BONAP county map]
Discussion

Physaria obdeltata is known from the eastern Snake River Plain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 635. FNA vol. 7, p. 654.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella Lesquerella obdeltata
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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