Physaria engelmannii |
Physaria navajoensis |
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Engelmann's bladderpod |
Navajo bladderpod |
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Habit | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). | Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). |
Stems | few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
several from base (crowded), erect, not exceeding leaves. |
Basal leaves | blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
usually absent. |
Cauline leaves | (proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
(petiole not differentiated from blade); blade linear-oblanceolate, 3–8(–13) mm, margins entire. |
Racemes | dense, (subumbellate). |
(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). |
Flowers | sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle). |
Fruiting pedicels | (ascending), relatively short. |
(ascending to divaricate-ascending, straight), 3.5–6 mm. |
Fruits | (shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4–8 per ovary; style 1.8–3 mm. |
Seeds | flattened. |
plump or slightly flattened, (strongly mucilaginous). |
2n | = 12, 24, 36. |
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Physaria engelmannii |
Physaria navajoensis |
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Phenology | Flowering Apr–May. | Flowering May-early Jun. |
Habitat | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops | Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone |
Elevation | 150-400 m (500-1300 ft) | 2200-2400 m (7200-7900 ft) |
Distribution |
OK; TX |
AZ; NM |
Discussion | Of conservation concern. Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 635. | FNA vol. 7, p. 651. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | Lesquerella navajoensis |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |