Physaria engelmannii |
Physaria ludoviciana |
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Engelmann's bladderpod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
Basal leaves | blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
Cauline leaves | (proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
Racemes | dense, (subumbellate). |
compact, (elongated and loose in fruit, densely-flowered). |
Flowers | sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
Fruiting pedicels | (ascending), relatively short. |
(usually recurved), (5–)10–20(–25) mm. |
Fruits | (shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
Seeds | flattened. |
slightly flattened. |
2n | = 12, 24, 36. |
= 10, 20, 30. |
Physaria engelmannii |
Physaria ludoviciana |
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Phenology | Flowering Apr–May. | Flowering Apr–Jul(-Aug). |
Habitat | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands |
Elevation | 150-400 m (500-1300 ft) | 0-1900 m (0-6200 ft) |
Distribution |
OK; TX |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 635. | FNA vol. 7, p. 649. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |