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Devils Gate twinpod

strong bladderpod

Habit Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). Perennials; caudex branched, (thickened); densely pubescent, trichomes (sessile), several-rayed, rays furcate or bifurcate, usually fused toward base, (strongly tuberculate).
Stems

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

several from base, erect or outer decumbent, (unbranched), to 2 dm.

Basal leaves

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

blade elliptic to lanceolate or obovate, 3–8 cm, margins entire.

Cauline leaves

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

(proximal shortly petiolate or sessile, distal sessile);

blade elliptic or obovate, to 2 cm, margins entire.

Racemes

condensed.

dense.

Flowers

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

sepals narrowly elliptic or oblong, 4.5–5.3 mm, (tapering to the somewhat thickened, cucullate apex, lateral pair subsaccate);

petals (bright yellow), lingulate or broadly obovate, 7.5–8.5 mm, (narrowing to broad claw, joining in an arch, margins lacerate).

Fruiting pedicels

(divaricate-ascending, nearly straight), 6–10 mm.

(divaricate-ascending to horizontal, straight to loosely curved), to 15 mm.

Fruits

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

(sessile or substipitate), suborbicular to broadly ovate or ellipsoid, slightly compressed, 6–8 mm;

valves pubescent;

ovules 12–22 per ovary;

style 2–3 mm.

Seeds

plump.

flattened.

2n

= 10.

Physaria eburniflora

Physaria valida

Phenology Flowering May–Jun. Flowering Apr–May.
Habitat Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus Limestone soils, steep slopes, roadcuts, open woods
Elevation 1800-3000 m (5900-9800 ft) 1900-2200 m (6200-7200 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
NM; TX
[BONAP county map]
Discussion

Physaria valida is known from the Sacramento and White mountains of south central New Mexico, and southward through the Guadalupe Mountains to Hudspeth County, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 635. FNA vol. 7, p. 664.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. vicina, P. vitulifera
Synonyms Lesquerella valida, Lesquerella lepidota
Name authority Rollins: Brittonia 33: 333. (1981) (Greene) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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