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Devils Gate twinpod

sessile bladderpod

Habit Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays usually furcate, rarely bifurcate, (moderately tuberculate or nearly smooth).
Stems

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

simple or few to several from base, erect, (often distal 1/2 branched), to 6 dm.

Basal leaves

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

blade oblanceolate, to 9 cm, margins entire, dentate, or sinuate to lyrate-pinnatifid.

Cauline leaves

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

(proximal shortly petiolate, distal sessile);

blade narrowly elliptic to linear, 2–4(–6) cm, margins entire or repand to shallowly dentate.

Racemes

condensed.

loose.

Flowers

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

sepals elliptic or elongate-ovate, 3.4–5.2(–6.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals obovate or deltate, 5–10 mm, (sometimes with short claw, margins undulate).

Fruiting pedicels

(divaricate-ascending, nearly straight), 6–10 mm.

(divaricate-ascending to widely spreading, straight), 8–20 mm.

Fruits

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

globose or subglobose, sometimes slightly compressed, 3–6 mm;

valves (not retaining seeds after dehiscence), densely papillose, densely pubescent inside, trichomes raised;

replum as wide as or wider than fruit;

ovules 8–18 per ovary;

style 1.5–3.5 mm.

Seeds

plump.

flattened.

2n

= 12.

Physaria eburniflora

Physaria sessilis

Phenology Flowering May–Jun. Flowering and fruiting Apr–Jun.
Habitat Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus Limestone chip, black soils, grassy roadsides, fields, limestone, oak woodlands, mesquite brush lands, pastures, open dry hills
Elevation 1800-3000 m (5900-9800 ft) 30-700 m (100-2300 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
TX
[BONAP county map]
Source FNA vol. 7, p. 635. FNA vol. 7, p. 663.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella gracilis var. sessilis, Lesquerella sessilis
Name authority Rollins: Brittonia 33: 333. (1981) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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