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Devils Gate twinpod

alpine bladderpod, reed's twinpod, Rollins' bladderpod

Habit Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout).
Stems

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm.

Basal leaves

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

(erect);

blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire.

Cauline leaves

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

blade linear, similar to basal.

Racemes

condensed.

dense, (often subumbellate, not or barely exceeding basal leaves).

Flowers

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals lingulate, 6–9 mm.

Fruiting pedicels

(divaricate-ascending, nearly straight), 6–10 mm.

(ascending, curved), 3–5.5 mm.

Fruits

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm;

valves pubescent, trichomes closely appressed;

ovules 8–12 per ovary;

style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma).

Seeds

plump.

plump, (oblong).

Physaria eburniflora

Physaria reediana

Phenology Flowering May–Jun. Flowering May–Jul.
Habitat Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus Open areas of grasslands on calcareous soils
Elevation 1800-3000 m (5900-9800 ft) 1200-1900 m (3900-6200 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
CO; NE; WY
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 635. FNA vol. 7, p. 660.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis
Name authority Rollins: Brittonia 33: 333. (1981) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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