Physaria douglasii |
Physaria thamnophila |
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Columbia bladder-pod, Douglas' bladder-pod |
Zapata bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex simple or branched, (woody); densely (silvery) pubescent, trichomes (short-stalked), 4–8-rayed, rays furcate or bifurcate, (finely tuberculate throughout). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
simple or few from base, decumbent, (straggling and flexuous, usually branched distally), 4–8 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade narrowly elliptic to oblanceolate, 4–12 cm, margins entire, sinuate, or shallowly dentate, (apex acute). |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(sessile or proximal shortly petiolate); blade linear to narrowly elliptic, 3–4 cm, margins entire, sinuate, or remotely dentate, (apex acute). |
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Racemes | loose (lax). |
loose, (sometimes greatly elongated). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals elliptic, 3.5–4 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals broadly obovate, 4–5 mm, (sometimes with short, broad claw). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(recurved), 15–20(–25) mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(pendent), subglobose or broadly ovoid, slightly compressed, 5–7 mm; valves (not retaining seeds after dehiscence), glabrous throughout; ovules per ovary unknown; style 1.5–2 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 10, 30. |
= 16. |
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Physaria douglasii |
Physaria thamnophila |
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Phenology | Flowering Apr. | |||||
Habitat | Sandy soils, entangled in shrubs, cactus clumps | |||||
Elevation | 1700-1800 m (5600-5900 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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TX |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria thamnophila is found in sandy areas with shrubs and cactus in sparse shrubland communities of Starr and Zapata counties. It sometimes flowers through September with sufficient moisture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 664. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Lesquerella thamnophila | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | ||||
Web links |
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