Physaria douglasii |
Physaria reediana |
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Columbia bladder-pod, Douglas' bladder-pod |
alpine bladderpod, reed's twinpod, Rollins' bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
(erect); blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
blade linear, similar to basal. |
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Racemes | loose (lax). |
dense, (often subumbellate, not or barely exceeding basal leaves). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals lingulate, 6–9 mm. |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(ascending, curved), 3–5.5 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm; valves pubescent, trichomes closely appressed; ovules 8–12 per ovary; style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma). |
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Seeds | flattened. |
plump, (oblong). |
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2n | = 10, 30. |
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Physaria douglasii |
Physaria reediana |
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Phenology | Flowering May–Jul. | |||||
Habitat | Open areas of grasslands on calcareous soils | |||||
Elevation | 1200-1900 m (3900-6200 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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CO; NE; WY
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 660. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||
Web links |
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