Physaria douglasii |
Physaria rectipes |
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Columbia bladder-pod, Douglas' bladder-pod |
straight bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; (loose, spreading); caudex simple or branched; densely pubescent, trichomes (subsessile), 4–6-rayed, rays furcate or bifurcate, (moderately tuberculate over arms, less so or smooth over center). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
few to several from base, ascending or prostrate, (arising laterally, also from within basal leaves, usually unbranched, rarely branched), 0.5–3(–6) dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade narrowly oblanceolate or broadly elliptic, 1–7(–12) cm, margins entire or shallowly toothed, sometimes repand, (inner blades usually flattened in age, surfaces often gray-green, scabrous). |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(usually secund); blade spatulate or obovate, 1–2.5(–4.5) cm, margins entire or shallowly toothed, (flat or involute). |
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Racemes | loose (lax). |
somewhat crowded (to moderately elongated in fruit, exceeding basal leaves). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals broadly elliptic or oblong, 4–7.5(–9) mm, (median pair thickened apically, cucullate); petals cuneate or obovate, 7–10(–16) mm, (tapering gradually to broad claw). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(often divaricate-spreading and straight, or horizontal and loosely sigmoid, sometimes slightly recurved), 5–15 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
subglobose to ovoid or ellipsoid, sometimes compressed, (4–)5–7(–9) mm; valves sparsely pubescent, trichomes appressed or erect, sometimes sparsely pubescent inside; ovules (8–)12–16(–20) per ovary; style 2–7 mm. |
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Seeds | flattened. |
somewhat flattened. |
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2n | = 10, 30. |
= 10 + 2, 18, 20, ± 40. |
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Physaria douglasii |
Physaria rectipes |
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Phenology | Flowering May–Jul. | |||||
Habitat | Sandy soils, limey knolls, rocky hills, clay hillsides, dry ridges, weathered rocks, gravelly outwashes, stony slopes, pinyon-juniper woodlands | |||||
Elevation | 1500-2600 m (4900-8500 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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AZ; CO; NM; UT
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As here circumscribed, Physaria rectipes remains heterogeneous and may represent more than one taxon, even after the recent removal of P. pulvinata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 660. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Lesquerella rectipes | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||
Web links |
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