Physaria douglasii |
Physaria mcvaughiana |
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Columbia bladder-pod, Douglas' bladder-pod |
Mcvaugh's bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex simple or branched, (sometimes enlarged); densely pubescent, trichomes (sessile), several-rayed, rays fused (webbed) most of their length, (umbonate, peltate, tuberculate throughout). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
few to several from base, erect or outer ones decumbent, 0.5–4 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
(long-petiolate); blade elliptic to obovate or rhombic, 2–6(–9) cm, margins entire. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(sessile or shortly petiolate); blade oblanceolate to spatulate, 1–3 cm, (proximal broader), margins entire. |
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Racemes | loose (lax). |
dense, (relatively short). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals elliptic or narrowly oblong, 4–5.4 mm, (tapered to apex); petals (white, base and claw yellow, conspicuously purple-veined), usually broadly obovate or rhombic, 6–10 mm, (± equal to blade, tapering to slender claw). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(erect to spreading, ascending, or (proximal) horizontal, straight to slightly curved, sometimes loosely sigmoid), 6–12(–20) mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(sessile or substipitate, often reddish magenta), usually ovoid to subglobose, inflated, 4–6(–7) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; septum perforate; ovules 8–12 per ovary; style 1.5–4 mm. |
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Seeds | flattened. |
somewhat flattened. |
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2n | = 10, 30. |
= 12. |
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Physaria douglasii |
Physaria mcvaughiana |
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Phenology | Flowering mid Mar–Apr(-Aug). | |||||
Habitat | Stream bed gravels, rocky limestone slopes and hills, canyon bottoms and slopes, limestone rubble | |||||
Elevation | 1200-1600 m (3900-5200 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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TX; Mexico (Coahuila) |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 650. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Lesquerella mcvaughiana | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||
Web links |
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