Physaria douglasii |
Physaria lindheimeri |
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Columbia bladder-pod, Douglas' bladder-pod |
Lindheimer's bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade pinnatisect to repand, 3–9(–14) cm, margins entire. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(sometimes secund, proximal usually petiolate, distal sessile); blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate. |
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Racemes | loose (lax). |
dense. |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate); petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth); valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules (8–)12–16(–20) per ovary; style (1.5–)2–3(–4) mm. |
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Seeds | flattened. |
flattened. |
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2n | = 10, 30. |
= 12. |
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Physaria douglasii |
Physaria lindheimeri |
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Phenology | Flowering Dec–Apr. | |||||
Habitat | Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies | |||||
Elevation | 20-800 m (100-2600 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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TX; Mexico (Tamaulipas) |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 649. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||
Web links |
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