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Columbia bladder-pod, Douglas' bladder-pod

point-tip twinpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Perennials; caudex branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays mostly furcate, (arms of unequal lengths, finely tuberculate).
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

several from base, erect or lateral decumbent, (unbranched), 1–2 dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

(petiole usually winged);

blade broadly oblanceolate, 3–8 cm, margins usually dentate or pinnatifid, rarely subentire, (terminal lobe acute or obtuse, not rounded).

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

blade spatulate to linear-oblanceolate, 1–3 cm, margins usually entire, rarely toothed, (apex acute).

Racemes

loose (lax).

loose (and greatly elongated in fruit) to congested.

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals linear-oblong, 5–7 mm;

petals spatulate, 9–11 mm.

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(recurved), 6–15 mm.

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

(usually pendent on arching pedicels, less frequently widely divergent), irregular in shape, (base obtuse or slightly cordate, apex deeply and broadly notched), not strongly inflated, 8–11 × 8–12 mm, (papery);

valves retaining seeds after dehiscence;

replum linear-oblong, constricted, 2.5–4 mm, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–8 mm.

Seeds

flattened.

flattened.

2n

= 10, 30.

Physaria douglasii

Physaria floribunda

Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
1. Fruit valves inflated, membranous; trichomes appressed.
subsp. floribunda
1. Fruit valves usually not inflated, coriaceous; trichomes slightly spreading.
subsp. osterhoutii
Source FNA vol. 7, p. 634. FNA vol. 7, p. 637.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
P. floribunda subsp. floribunda, P. floribunda subsp. osterhoutii
Synonyms Lesquerella douglasii
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Rydberg: Bull. Torrey Bot. Club 28: 279. (1901)
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