Physaria douglasii |
Physaria filiformis |
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Columbia bladder-pod, Douglas' bladder-pod |
limestone glade or Missouri bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Annuals; with a fine taproot; densely pubescent, trichomes (sessile), usually 4-rayed, rays forked, rarely simple or tripartite, (finely tuberculate). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
several from base, (slender), erect or outer decumbent, (usually branched, branches filiform, bud clusters of growing plants drooping), to 2.5 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade orbicular to broadly spatulate, 1–2.4 cm, margins entire or sinuate. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(proximal often petiolate, distal sessile); similar to basal, blade spatulate to oblanceolate or (distal) linear, (base cuneate), margins entire or sinuate. |
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Racemes | loose (lax). |
loose. |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals oblong or elliptic, 2.5–4.6 mm, (median pair slightly thickened apically); petals (pale yellow), spatulate to obovate, 5–9 mm, (apex ± emarginate). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(usually divaricate-ascending, straight or slightly curved), 7–11 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(sessile or shortly stipitate), globose, not inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 3–5 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 10, 30. |
= 14. |
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Physaria douglasii |
Physaria filiformis |
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Phenology | Flowering Apr–May. | |||||
Habitat | Limestone, dolomite, and shale, sparsely vegetated or barren areas, cedar glades, old pastures, along roadsides | |||||
Elevation | 200-300 m (700-1000 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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AL; AR; MO |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 637. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Lesquerella filiformis | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |
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