Physaria douglasii |
Physaria carinata |
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Columbia bladder-pod, Douglas' bladder-pod |
Idaho bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). | ||||||||||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(sessile or shortly petiolate); blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire. |
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Racemes | loose (lax). |
compact to loose. |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not); petals spatulate, 7.5–10 mm. |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled); valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside; ovules (4–)8–14(–16) per ovary; style 2–4.5(–5) mm. |
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Seeds | flattened. |
slightly flattened. |
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2n | = 10, 30. |
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Physaria douglasii |
Physaria carinata |
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Distribution |
ID; MT; OR; WA; BC
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ID; MT; WY |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 629. | ||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Lesquerella douglasii | Lesquerella carinata | ||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||||||||||
Web links |
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