Physaria douglasii |
Physaria calderi |
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Columbia bladder-pod, Douglas' bladder-pod |
Calder's bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade oblanceolate, 2–3 cm, margins entire. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
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Racemes | loose (lax). |
loose. |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
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Seeds | flattened. |
plump. |
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2n | = 10, 30. |
= 20. |
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Physaria douglasii |
Physaria calderi |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Dry rocky summits, limestone flats and slopes, alpine knolls | |||||
Elevation | 600-1500 m (2000-4900 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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AK; NT; YT |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 629. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Lesquerella calderi, Lesquerella arctica subsp. calderi | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
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