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Columbia bladder-pod, Douglas' bladder-pod

Great Plains bladderpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate).
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat).

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire.

Racemes

loose (lax).

(secund), dense, (elongated in fruit).

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw).

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout).

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm;

valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside;

ovules (4–)8(–10) per ovary;

style (slender), 3–5.5(–6.5) mm.

Seeds

flattened.

slightly flattened.

2n

= 10, 30.

Physaria douglasii

Physaria arenosa

Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; ND; NE; SD; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
1. Perennials or, rarely, annuals, short-lived; fruit valves: trichomes spreading.
subsp. arenosa
1. Perennials, long-lived; fruit valves: trichomes closely appressed.
subsp. argillosa
Source FNA vol. 7, p. 634. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
P. arenosa subsp. arenosa, P. arenosa subsp. argillosa
Synonyms Lesquerella douglasii Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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