Physaria douglasii |
Physaria acutifolia |
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Columbia bladder-pod, Douglas' bladder-pod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
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Racemes | loose (lax). |
loose, (elongated in fruit). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
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Seeds | flattened. |
(dark brown), flattened, (2–3 mm). |
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2n | = 10, 30. |
= 10, 16, 24. |
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Physaria douglasii |
Physaria acutifolia |
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Phenology | Flowering May–Jun(-Jul). | |||||
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | |||||
Elevation | 1500-3500 m (4900-11500 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 624. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | ||||
Web links |
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