Physaria douglasii |
Brassicaceae tribe Physarieae |
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Columbia bladder-pod, Douglas' bladder-pod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Annuals, biennials, perennials, or subshrubs; eglandular. | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
petiolate, sessile, or subsessile; blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate. |
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Racemes | loose (lax). |
ebracteate, often elongated in fruit. |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
actinomorphic; sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally; petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct; filaments unappendaged, not winged; pollen (3 or) 4–11-colpate. |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate; ovules 2–100 per ovary; style usually distinct; stigma entire or strongly 2-lobed. |
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Seeds | flattened. |
biseriate, uniseriate, or aseriate; cotyledons accumbent or incumbent. |
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Trichomes | usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones. |
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2n | = 10, 30. |
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Physaria douglasii |
Brassicaceae tribe Physarieae |
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Distribution |
ID; MT; OR; WA; BC
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North America; Mexico; South America; Asia (ne Russia) |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 7, species ca. 130 (7 genera, 105 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 604. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | |||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895) | ||||
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