Physaria didymocarpa |
Physaria pycnantha |
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common twinpod, double twinpod, twin bladderpod |
mountain-view bladderpod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex branched, (densely cespitose and forming hemispheric mounds); densely pubescent, trichomes 5-rayed, rays bifurcate near base, fused at base, (strongly tuberculate throughout). | ||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
few to several from base, erect, (usually exceeding basal leaves), 0.3–0.7 dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
blade linear-spatulate, 1.5–4 cm, (base narrowed gradually to petiole), margins entire. |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
blade spatulate, similar to basal. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
crowded in distal 1/3, (4–10-flowered). |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals (pale yellow), oblong to elliptic, 3–4 mm, (median pair usually thickened apically, cucullate); petals (sometimes with slight tinge of orange basally), lingulate, 4–6 mm. |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(loosely to strongly sigmoid), 6–10 mm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
ellipsoid, slightly inflated (somewhat latiseptate), 4–5 mm, (apex acute); valves pubescent, trichomes erect, appearing slightly shaggy; ovules 4–8 per ovary; styles 2.5–3 mm, (shorter than mature fruits). |
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Seeds | flattened. |
± flattened, convex on outer side. |
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Physaria didymocarpa |
Physaria pycnantha |
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Phenology | Flowering late May–Jun(-Jul). | |||||||||
Habitat | Dry, windswept knolls of limestone gravel, with other cushion-forming plants | |||||||||
Elevation | 1600-2300 m (5200-7500 ft) | |||||||||
Distribution |
ID; MT; WA; WY; AB; BC
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ID; MT
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pycnantha is morphologically similar to 56. P. nelsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 659. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | |||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | Grady & O’Kane: Novon 17: 188, fig. 5. (2007) | ||||||||
Web links |