Physaria didymocarpa |
Physaria purpurea |
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common twinpod, double twinpod, twin bladderpod |
rose bladderpod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). | ||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
simple from base, erect, (unbranched, sparsely leaved), to 7 dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid. |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
(proximal often narrowed to short petiole, distal sessile); blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
dense or slightly elongated. |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate); petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade). |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(spreading or recurved, loosely sigmoid), 5–25 mm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
(pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 1–3(–4) mm. |
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Seeds | flattened. |
flattened. |
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2n | = 18, 36. |
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Physaria didymocarpa |
Physaria purpurea |
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Phenology | Flowering Mar–Oct. | |||||||||
Habitat | Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps | |||||||||
Elevation | 400-2400 m (1300-7900 ft) | |||||||||
Distribution |
ID; MT; WA; WY; AB; BC
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AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 659. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa | ||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||||||
Web links |