Physaria didymocarpa |
Physaria prostrata |
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common twinpod, double twinpod, twin bladderpod |
low bladderpod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex branched; densely pubescent, trichomes (usually sessile, rarely short-stalked), several-rayed, rays furcate or bifurcate, (umbonate, tuberculate throughout). | ||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
several from base, usually prostrate, rarely decumbent, (unbranched, often purplish, sparsely pubescent), to 1.5 dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
blade deltate, hastate, or, less often, rhombic to elliptic, 1–5 cm, margins entire (often partially involute). |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
(proximal shortly petiolate); blade linear to oblanceolate, 0.5–1.5 cm, margins entire. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
loose, elongated. |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals (often purplish), oblong, lanceolate, or ovate, 4–6 mm, (median pair thickened apically); petals spatulate or cuneate, 5–8(–9) mm, (margins undulate). |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(ascending, slightly sigmoid to straight), 5–10 mm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
ovoid or ellipsoid, slightly compressed, 3–5(–6) mm, (base often gibbous); valves pubescent, trichomes loose, furcate near their bases and spreading, sometimes sparsely pubescent inside; ovules 4(–8) per ovary; style 1.5–4 mm. |
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Seeds | flattened. |
flattened. |
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Physaria didymocarpa |
Physaria prostrata |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Whitish sand and small rocks on steep slopes, dry hillsides, windswept knolls, shaley slopes | |||||||||
Elevation | 1800-2500 m (5900-8200 ft) | |||||||||
Distribution |
ID; MT; WA; WY; AB; BC
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ID; UT; WY |
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria prostrata is sometimes found on igneous substrates, which is unusual for the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 658. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Lesquerella prostrata | ||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||||||
Web links |