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common twinpod, double twinpod, twin bladderpod

low bladderpod

Habit Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). Perennials; caudex branched; densely pubescent, trichomes (usually sessile, rarely short-stalked), several-rayed, rays furcate or bifurcate, (umbonate, tuberculate throughout).
Stems

several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm.

several from base, usually prostrate, rarely decumbent, (unbranched, often purplish, sparsely pubescent), to 1.5 dm.

Basal leaves

(forming a strong rosette; long-petiolate);

blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery).

blade deltate, hastate, or, less often, rhombic to elliptic, 1–5 cm, margins entire (often partially involute).

Cauline leaves

blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute).

(proximal shortly petiolate);

blade linear to oblanceolate, 0.5–1.5 cm, margins entire.

Racemes

congested, (elongated in fruit, greatly exceeding leaves).

loose, elongated.

Flowers

sepals lanceolate to oblong, 6–8 mm, (often keeled);

petals spatulate, 10–12 mm.

sepals (often purplish), oblong, lanceolate, or ovate, 4–6 mm, (median pair thickened apically);

petals spatulate or cuneate, 5–8(–9) mm, (margins undulate).

Fruiting pedicels

(spreading, straight or slightly curved), 8–12 mm.

(ascending, slightly sigmoid to straight), 5–10 mm.

Fruits

(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy);

replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse;

ovules (4–)8 per ovary;

style 7–9 mm.

ovoid or ellipsoid, slightly compressed, 3–5(–6) mm, (base often gibbous);

valves pubescent, trichomes loose, furcate near their bases and spreading, sometimes sparsely pubescent inside;

ovules 4(–8) per ovary;

style 1.5–4 mm.

Seeds

flattened.

flattened.

Physaria didymocarpa

Physaria prostrata

Phenology Flowering May–Jun.
Habitat Whitish sand and small rocks on steep slopes, dry hillsides, windswept knolls, shaley slopes
Elevation 1800-2500 m (5900-8200 ft)
Distribution
from FNA
ID; MT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; UT; WY
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria prostrata is sometimes found on igneous substrates, which is unusual for the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trichomes spreading throughout; basal leaf blade margins dentate.
subsp. lanata
1. Trichomes appressed (except fruits in subsp. didymocarpa); basal leaf blade margins repand to dentate, ± lyrate or, rarely, entire
→ 2
2. Fruits moderately inflated, trichomes spreading.
subsp. didymocarpa
2. Fruits highly inflated (papery), trichomes appressed.
subsp. lyrata
Source FNA vol. 7, p. 633. FNA vol. 7, p. 658.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. didymocarpa subsp. didymocarpa, P. didymocarpa subsp. lanata, P. didymocarpa subsp. lyrata
Synonyms Vesicaria didymocarpa, Coulterina didymocarpa Lesquerella prostrata
Name authority (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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