Physaria didymocarpa |
Physaria montana |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
common twinpod, double twinpod, twin bladderpod |
mountain bladderpod |
|||||||||
Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex simple or branched, (often enlarged); densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). | ||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
simple or several from base, prostrate to erect, 0.5–2(–3.5) dm. |
||||||||
Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
blade suborbicular or obovate to elliptic, (1–)2–5(–7) cm, margins entire, sinuate, or shallowly dentate. |
||||||||
Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
(often secund, proximal shortly petiolate, distal sessile); blade linear to obovate or rhombic, 1–2.5(–4) cm, margins entire or shallowly dentate. |
||||||||
Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
dense, compact, (usually elongated in fruit). |
||||||||
Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals elliptic, 5–8.5 mm, (lateral pair boat-shaped, saccate, median pair thickened apically, cucullate); petals (yellow to orange, sometimes fading purplish), narrowly spatulate or obovate, (6–)7.5–12 mm, (claw undifferentiated from blade, or gradually narrowed to claw, slightly expanded basally). |
||||||||
Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(usually sharply sigmoid, rarely nearly divaricate-spreading and straight), 5–15(–20) mm, (stout). |
||||||||
Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
(erect), ellipsoid or ovoid, not or slightly obcompressed, (apex not compressed), (6–)7–12 mm; valves densely pubescent, sometimes sparsely pubescent inside; ovules (8–)12–20(–24) per ovary; style 3–7 mm, (sometimes pubescent). |
||||||||
Seeds | flattened. |
flattened. |
||||||||
2n | = 10. |
|||||||||
Physaria didymocarpa |
Physaria montana |
|||||||||
Phenology | Flowering Apr–Jun(-Aug). | |||||||||
Habitat | Banks, rock outcrops, stony slopes and benchlands, from plains into mountains, in sagebrush, open scrub oak, pinyon-juniper woodland, ponderosa pine, Douglas fir on granitic, often gravelly, non-calcareous soils, rarely on calcareous soils | |||||||||
Elevation | 1000-3300 m (3300-10800 ft) | |||||||||
Distribution |
ID; MT; WA; WY; AB; BC
|
AZ; CO; NE; NM; SD; WY
|
||||||||
Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria montana is a rather variable species that in southwestern Colorado morphologically approaches P. rectipes and in eastern Wyoming approaches P. curvipes; it is unusual in the genus for its frequent presence on igneous, non-calcareous soils. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||
Key |
|
|||||||||
Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 650. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Vesicaria montana, Alyssum grayanum, Lesquerella montana, Lesquerella montana var. suffruticosa, Lesquerella rosulata | ||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (A. Gray) Greene: Fl. Francisc., 249. (1891) | ||||||||
Web links |