Physaria didymocarpa |
Physaria geyeri |
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common twinpod, double twinpod, twin bladderpod |
Geyer's bladderpod, Geyer's twin-pod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). | ||||||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
several from base, decumbent, (arising laterally, unbranched), 1–3 dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
(numerous); (petiole slender, rarely with a few broad teeth); blade obovate, 3–7 cm, margins entire. |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
blade oblanceolate, 1.5–3 cm, margins entire. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
loose. |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals oblong, 5–7 mm; petals (yellow to purplish), spatulate, 8–12 mm. |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(ascending or spreading, slightly curved or sigmoid), 1–2 cm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading; replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse; ovules 4–6 per ovary; style 5–7 mm. |
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Seeds | flattened. |
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Physaria didymocarpa |
Physaria geyeri |
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Distribution |
ID; MT; WA; WY; AB; BC
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ID; MT; OR; WA
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 638. | ||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri | ||||||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||||||
Web links |