Physaria didymocarpa |
Physaria fremontii |
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common twinpod, double twinpod, twin bladderpod |
Fremont's bladderpod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex simple; densely pubescent, trichomes (subsessile), 5–7-rayed, rays distinct or slightly fused at base, usually furcate, (roughly tuberculate). | ||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
few to several from base, prostrate, (arising proximal to a terminal cluster of erect leaves, usually unbranched, slender), 0.5–1(–1.5) dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
(petiole slender); blade elliptic to rhombic, 1.5–4(–5) cm, (base gradually tapering to petiole), margins usually entire, rarely dentate. |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
blade narrowly obovate to oblanceolate, 5–15 mm, margins entire. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
(relatively short), rather loose. |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals ± elliptic, 4–6 mm; petals ovate, 6–8 mm, (claw slightly expanded at base). |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(usually secund, recurved), 5–8 mm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
(pendent), globose, subglobose, or slightly obcompressed, slightly or not inflated, 3–6 mm, (rigid, apex usually beaked); valves pubescent throughout; ovules 8–12 per ovary; style 1–2 mm, (pubescent). |
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Seeds | flattened. |
slightly flattened, (suborbicular). |
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Physaria didymocarpa |
Physaria fremontii |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Calcareous gravel, loose whitish rubble, limestone pavement, rocky calcareous ridges | |||||||||
Elevation | 2100-2800 m (6900-9200 ft) | |||||||||
Distribution |
ID; MT; WA; WY; AB; BC
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WY |
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria fremontii is known from the area of the Wind River Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 638. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Lesquerella fremontii | ||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||||||
Web links |