Physaria didymocarpa |
Physaria carinata |
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common twinpod, double twinpod, twin bladderpod |
Idaho bladderpod |
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Habit | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate). | ||||||||||||||||
Stems | several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm. |
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Basal leaves | (forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed. |
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Cauline leaves | blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
(sessile or shortly petiolate); blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire. |
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Racemes | congested, (elongated in fruit, greatly exceeding leaves). |
compact to loose. |
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Flowers | sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not); petals spatulate, 7.5–10 mm. |
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Fruiting pedicels | (spreading, straight or slightly curved), 8–12 mm. |
(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm. |
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Fruits | (erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled); valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside; ovules (4–)8–14(–16) per ovary; style 2–4.5(–5) mm. |
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Seeds | flattened. |
slightly flattened. |
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Physaria didymocarpa |
Physaria carinata |
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Distribution |
ID; MT; WA; WY; AB; BC
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ID; MT; WY |
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 633. | FNA vol. 7, p. 629. | ||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | Lesquerella carinata | ||||||||||||||||
Name authority | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||||||||||||||
Web links |