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denseflower bladderpod, low bladderpod

manyhead bladderpod

Habit Annuals or biennials; caudex simple or branched, (relatively small, cespitose); densely pubescent, trichomes (spreading, sessile or short-stalked), 5–7-rayed, rays distinct and simple, (tuberculate, finely tubercled with a U-shaped notch on one side). Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout).
Stems

simple or few to several from base, erect or decumbent, (rarely branched, usually leafy), to 4 dm.

several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm.

Basal leaves

blade lyrate-pinnatifid, 1–7 cm, margins entire or shallowly dentate.

blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery).

Cauline leaves

(sessile or shortly petiolate);

blade narrowly obovate to elliptic, 0.5–6 cm, margins entire, repand, or shallowly dentate.

blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent).

Racemes

dense, (elongated in fruit, often subtended by distal cauline leaves).

(narrow), loose, (elongated in fruit).

Flowers

sepals elliptic, 3.7–7.2 mm, (lateral pair somewhat cucullate, median pair thickened apically);

petals (yellow to orange-yellow), obovate to obdeltate, (4.5–)7–10(–11) mm, (tapering to short claw, apex often emarginate).

sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate);

petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade).

Fruiting pedicels

(usually divaricate-spreading, straight or slightly curved, delicate, sometimes drooping, especially on herbarium specimens), 7–10 mm, (somewhat rigid).

(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm.

Fruits

(sessile or substipitate), globose or broadly obovate, not inflated, 4–6 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 2–5 mm.

broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm;

valves sparsely pubescent;

ovules usually 4, rarely 6–8 per ovary;

style 3–6.5 mm.

Seeds

flattened.

plump.

2n

= 14.

Physaria densiflora

Physaria multiceps

Phenology Flowering Mar–May. Flowering May–Jul.
Habitat Sandy, granitic, or calcareous soils, sandy ledges, limestone outcrops, rocky prairies, uplands Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks
Elevation 30-400 m (100-1300 ft) 2400-2900 m (7900-9500 ft)
Distribution
from FNA
TX
[BONAP county map]
from FNA
ID; UT; WY
[BONAP county map]
Discussion

Alyssum densiflorum (A. Gray) Kuntze (1891), not Desfontaines (1808) is an illegitimate name, sometimes found in synonymy with Physaria densiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 633. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria densiflora, Lesquerella densiflora Lesquerella multiceps
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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