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denseflower bladderpod, low bladderpod

Devils Gate twinpod

Habit Annuals or biennials; caudex simple or branched, (relatively small, cespitose); densely pubescent, trichomes (spreading, sessile or short-stalked), 5–7-rayed, rays distinct and simple, (tuberculate, finely tubercled with a U-shaped notch on one side). Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth).
Stems

simple or few to several from base, erect or decumbent, (rarely branched, usually leafy), to 4 dm.

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

Basal leaves

blade lyrate-pinnatifid, 1–7 cm, margins entire or shallowly dentate.

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

Cauline leaves

(sessile or shortly petiolate);

blade narrowly obovate to elliptic, 0.5–6 cm, margins entire, repand, or shallowly dentate.

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

Racemes

dense, (elongated in fruit, often subtended by distal cauline leaves).

condensed.

Flowers

sepals elliptic, 3.7–7.2 mm, (lateral pair somewhat cucullate, median pair thickened apically);

petals (yellow to orange-yellow), obovate to obdeltate, (4.5–)7–10(–11) mm, (tapering to short claw, apex often emarginate).

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

Fruiting pedicels

(usually divaricate-spreading, straight or slightly curved, delicate, sometimes drooping, especially on herbarium specimens), 7–10 mm, (somewhat rigid).

(divaricate-ascending, nearly straight), 6–10 mm.

Fruits

(sessile or substipitate), globose or broadly obovate, not inflated, 4–6 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 2–5 mm.

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

Seeds

flattened.

plump.

2n

= 14.

Physaria densiflora

Physaria eburniflora

Phenology Flowering Mar–May. Flowering May–Jun.
Habitat Sandy, granitic, or calcareous soils, sandy ledges, limestone outcrops, rocky prairies, uplands Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus
Elevation 30-400 m (100-1300 ft) 1800-3000 m (5900-9800 ft)
Distribution
from FNA
TX
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Alyssum densiflorum (A. Gray) Kuntze (1891), not Desfontaines (1808) is an illegitimate name, sometimes found in synonymy with Physaria densiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 633. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria densiflora, Lesquerella densiflora
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Rollins: Brittonia 33: 333. (1981)
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