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tuft twinpod

alpine bladderpod, spatula-leaf bladderpod

Habit Perennials; caudex usually simple, rarely branched, (enlarged with persistent leaf bases, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays typically furcate, (fused at base, arms slender, tuberculate throughout). Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate).
Stems

several from base, decumbent to ascending, (arising laterally beneath a dense rosette), less than 0.1 dm.

simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm.

Basal leaves

(petiole slender);

blade (horizontal on the ground), obovate, 0.5–1.5 cm (width 4–8 mm, base tapering abruptly to petiole), margins entire, (apex usually acute, surfaces silvery from a dense incrustation of appressed, stellate trichomes).

(erect to prostrate, petiole distinct from blade);

blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded).

Cauline leaves

blade oblanceolate, 0.5–1 cm (width 2–3 mm), margins entire, (surfaces densely stellate pubescent).

blade spatulate, distinctly different from basal.

Racemes

congested, (subumbellate, often almost sessile, barely exceeding basal leaves).

moderately dense, (6–20-flowered).

Flowers

sepals (yellowish green), narrowly lanceolate, 4–5 mm;

petals (erect), oblanceolate, 6–7 mm, (claw weakly differentiated from blade).

sepals (pale yellow), elliptic, 3.5–5 mm;

petals lingulate, 6–9 mm.

Fruiting pedicels

(divaricate, straight), 5–10 mm.

(strongly sigmoid), 10–20 mm (2 times longer than fruits).

Fruits

didymous, ovate, inflated, 4.8–6 × 6–10 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes loosely spreading;

replum obovate, 3–4 mm, as wide as or wider than fruit;

ovules 8 per ovary (2–4 abortive);

style 4–6 mm.

lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed);

valves pubescent, trichomes sparse and closely appressed to surface;

ovules 4–8 per ovary;

style 2.5–6 mm (usually ± equal in length to mature fruit).

Seeds

flattened.

plump.

Physaria condensata

Physaria spatulata

Phenology Flowering May–Jun. Flowering May-early Jul.
Habitat Calcareous knolls and ridges, clay banks, limey slopes, shaley hills, clay patches Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates
Elevation 1800-2400 m (5900-7900 ft) 900-2900 m (3000-9500 ft)
Distribution
from FNA
WY
[BONAP county map]
from FNA
MT; ND; NE; SD; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 631. FNA vol. 7, p. 663.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata
Name authority Rollins: Rhodora 41: 407, plate 556, figs. 1, 9, 10. (1939) (Rydberg) Grady & O’Kane: Novon 17: 190. (2007)
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