Physaria chambersii |
Physaria tumulosa |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Kodachrome bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex (buried), branched, (forming hard mats); densely pubescent, trichomes several-rayed, rays furcate or bifurcate, (tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect, (unbranched), 0.2–0.3 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
(few), similar to cauline. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(petiole not differentiated from blade); blade (somewhat succulent), linear to narrowly oblanceolate, 5–12 mm, margins entire. |
Racemes | congested. |
dense, (few-flowered). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (yellowish), elliptic, 3–4.5 mm; petals (erect or, more commonly, arching), spatulate to oblanceolate, 5.8–7 mm, (claw not or weakly differentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(ascending to divaricate-ascending, ± straight), 3.5–6 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(coppery or reddish brown in age), broadly ovoid, slightly inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.8–3 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
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Physaria chambersii |
Physaria tumulosa |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Barren white knolls surrounded by sagebrush, pinyon pine, and Utah juniper |
Elevation | 1500-3200 m (4900-10500 ft) | 1600-1800 m (5200-5900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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UT |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria tumulosa is morphologically similar to 55. P. navajoensis of northeastern Arizona and northwestern New Mexico, and differing very subtly. It has been long treated as an infraspecific taxon of P. hitchcockii; unpublished molecular data do not support that disposition. It is found on knolls of the Winsor Member of the Carmel Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 664. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella hitchcockii subsp. tumulosa, Lesquerella tumulosa, P. rubicundula var. tumulosa |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Barneby) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
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