Physaria chambersii |
Physaria thamnophila |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
Zapata bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (woody); densely (silvery) pubescent, trichomes (short-stalked), 4–8-rayed, rays furcate or bifurcate, (finely tuberculate throughout). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
simple or few from base, decumbent, (straggling and flexuous, usually branched distally), 4–8 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade narrowly elliptic to oblanceolate, 4–12 cm, margins entire, sinuate, or shallowly dentate, (apex acute). |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(sessile or proximal shortly petiolate); blade linear to narrowly elliptic, 3–4 cm, margins entire, sinuate, or remotely dentate, (apex acute). |
Racemes | congested. |
loose, (sometimes greatly elongated). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals elliptic, 3.5–4 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals broadly obovate, 4–5 mm, (sometimes with short, broad claw). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(recurved), 15–20(–25) mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(pendent), subglobose or broadly ovoid, slightly compressed, 5–7 mm; valves (not retaining seeds after dehiscence), glabrous throughout; ovules per ovary unknown; style 1.5–2 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 16. |
Physaria chambersii |
Physaria thamnophila |
|
Phenology | Flowering Apr–Jul. | Flowering Apr. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Sandy soils, entangled in shrubs, cactus clumps |
Elevation | 1500-3200 m (4900-10500 ft) | 1700-1800 m (5600-5900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
|
TX |
Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria thamnophila is found in sandy areas with shrubs and cactus in sparse shrubland communities of Starr and Zapata counties. It sometimes flowers through September with sufficient moisture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 664. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella thamnophila |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
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