Physaria chambersii |
Physaria tenella |
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Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
little bladderpod, Moapa bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Annuals or, rarely, biennials; with a taproot; densely pubescent, trichomes (simple or stellate, sessile or short-stalked), 4–7-rayed, rays usually furcate, rarely bifurcate, (nearly smooth to finely tuberculate). |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, decumbent to erect, (several-branched, frequently stout), 1.5–6 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade elliptic, (1.5–)3–6.5 cm, margins entire, repand, or shallowly dentate. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
(proximal shortly petiolate, distal sessile); blade linear to elliptic or obovate, (0.5–)1–3.5(–4.5) cm, margins entire or repand. |
Racemes | congested. |
loose. |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals oblong, lanceolate, or elliptic, (3–)3.5–6(–7.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (yellow to orange), suborbicular or obovate, (5–)6.5–8(–11) mm, (narrowing gradually to broad claw, usually widened at base). |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(recurved, sigmoid), 5–15 mm. |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(sessile or shortly stipitate), orbicular or obovoid, often slightly compressed, (3.5–)4–6 mm; valves sparsely pubescent, trichomes sessile and stellate, densely pubescent inside, trichomes simple or branched; ovules 4–12 per ovary; style 2–4.5 mm. |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
= 10, 20. |
Physaria chambersii |
Physaria tenella |
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Phenology | Flowering Apr–Jul. | Flowering Feb–May. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Sandy soils, gravel, clayey loam, loose rocky slopes, washes, desert slopes and plains, lava hills, frequently in or near bushes |
Elevation | 1500-3200 m (4900-10500 ft) | (0-)600-1900 m ((0-)2000-6200 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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AZ; CA; NV; UT; Mexico (Sonora)
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella tenella, Lesquerella gordonii var. sessilis |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
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