Physaria chambersii |
Physaria subumbellata |
|
---|---|---|
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
parasol bladderpod |
|
Habit | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | Perennials; caudex simple or branched, (usually covered with persistent leaf bases, cespitose); densely pubescent, trichomes (closely appressed), rays distinct, usually bifurcate. |
Stems | several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
several from base, erect, (unbranched, slender), 0.1–0.6 dm. |
Basal leaves | (petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
blade rhombic to obovate, 2–4 cm, margins entire. |
Cauline leaves | blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
blade linear-oblanceolate, similar to basal. |
Racemes | congested. |
dense (distally, subumbellate). |
Flowers | sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
sepals (yellowish), oblong to elliptic, 3.5–7 mm, (median pair usually thickened apically, cucullate); petals lingulate to spatulate, 4–7 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid), 8–15 mm. |
(divaricate-ascending), 3–5 mm, (densely pubescent). |
Fruits | (often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
(erect), ovate to suborbicular, compressed apically (latiseptate), 3–4 mm; valves pubescent; replum ovate to obovate; ovules 4–6 per ovary; style 2–3 mm. |
Seeds | flattened. |
plump. |
2n | = 8, 10, 16, 24. |
= 10. |
Physaria chambersii |
Physaria subumbellata |
|
Phenology | Flowering Apr–Jul. | Flowering Apr–Jun. |
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | Rocky high ridges, gravel and stony areas, juniper covered knolls, rock crevices, clay hillsides, pinyon-juniper areas, calcareous substrates |
Elevation | 1500-3200 m (4900-10500 ft) | 1600-2700 m (5200-8900 ft) |
Distribution |
AZ; CA; ID; NV; OR; UT
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CO; UT; WY
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 631. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | Lesquerella subumbellata |
Name authority | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
Web links |